New and little known species of Dolichopodidae from China ...

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BULLE T! DE L' I NSTITUT ROYAL DES SC I ENCES NATURE LL ES DE BE LG I QUE E TOMOLOGI E. 70: 219-24 2. 20 00 EN TOMOL OG I E. 70: 21 9,- 242. 20 00 BU L LETI N VAN HET KON I NK LIJ K BELG I SCH I NSTI T UUT VOOR AT UU RWETENSCHA PI' EN New and little known species of Dolichopodidae from China (VII): Diptera fro1n Etnei Mountain (2) by DingY ANG and Toyohei SAIGUSA Abstract - 20 species of th e famil y Dolichopodidae from Emei Mountain are described as new to science: !-l ercos/omus (!-!.) curvarmalus, H. (!-!.) curvispinus. H. (H) incisus. 1-1. (!-!.) j indinganus, H (!-! .) longus. 1-1. (!-!.) 5piniwrsis, !-!. (H ) sub/angus, Dioslracus brevicercus, D. br evis. D. dig ilif ormi s, D. prolongalus. Amblypsilopus !ongisew. lv l edelera curvala, Diaphorus lridenla/us. Rlwphium Jitrcalunl , Neuri gonel!a !ongisew, N. pallipes. N. lri/itrcaw. Teuchoplwrus emeiensis, T. ven- /ralis. Key wor·ds: Dolic hopoclidae, new species, Emei Mountain, Sichuan Introduction II genera a nd 41 species of th e family Dolichopodidae are known from Emei Mounta in (YANG & SA I GUSA , 1999). Mainly bas ed on th e mate ri al co ll ected by th e second a uth or, we add here 29 species to thi s region. Among th em 20 species are described as new to science. The types of new species are mainly depos ited in the Ch in a Agr ic ultural Uni vers it y (CAU) except some para- types kept in th eSA I GU SA co ll ec ti on at the Kyushu Uni- versity (SKU) and the '' In stitut Royal des Sciences Nat- urelles de Belgique" (IRSN) . The fo llowing abbreviations are use d: acr-acrostichal, ad-anterodorsal, apv-apicoventral, av-anteroventral, d- dorsal, dc-dorsocentral, h-hume ral , ih - inn er humeral, U-fore leg, LII-mid leg, LIII-hind leg, np l-notopleural, oc-oce ll ar, pa- postalar, pd-posterodorsal, ph-posthum- eral, psu-postsutural, pv-posteroventral, pvt-postve rti cal, sa-supraalar, su-sutural, v-ve ntral, vt-ve rti ca l. S ubf am ily Do li cho podina e l. Plutlacrosoma briarea WEI et LJU Phalacrosoma briarea WEI et LIU , 1996. J. Guizhou Ag ri c. Co li. I 5( 1 ): 38 . Type locality: Ans hun , Guizhou. Sp ecimens examined. I male, Sic hu an: Emei Mountain, Maz ib a ( I, I 00- 1,300 m) , 1998. VIII. 16, T. SAIG US A; 3 ma les, Sichuan: Emei Mo un ta in, Jings hui (700-750 m) , 1998. VI II. 7 T. SAI GUSA; I male, Sichuan: Qingcheng Mountain, Yuechenghu (900 m), 1998. VIII. 22, T. SA I- GUSA (S KU). Distribution. China (Sichuan, Guizhou). 2. Hercos tomus (Hercostomus) curvispiuus sp. nov. (Figs. 1-2) Diagnosis. Postoc ul ar bristles wholly black. First tlage l- lomere sma ll an d acute apically. Fore tarsomeres 3-4 black and slightly thi ckened, with thi ck dorsal hairs; tarsomere 5 whi te with white ha irs. Ma l e. Body leng th 2 .8 mm , win g length 3.2 mm. Head meta lli c green with pale grey po ll en; vertex, frons and face with s il ve ry whi te po ll en; face nea rl y as wid e as first tlagellomere. Hairs and bristles on head bl ack; postoc ul ar bristles black, mid-ventral hai rs pale. Antenn a small and bl ack; first tlage ll omere as long as apica ll y; _arista black, with basal segment 0. 17 t1mes as long as ap1cal segment, very short pub escent. Proboscis ye ll ow with bl ack hairs; palpus ye ll ow, with black hairs and I black apical bristle. Thorax meta llic green with pale grey po ll en. Ha irs and bri s tl es on thorax bl ack; 6 strong de; 5-6 irregularl y paired acr short and hai r-like; scutellum with two pa irs of bris tl es (outer pair very short, hair-like). Propleuron pale haired, with I bl ack bristle on lower portion. Legs ye ll ow; fore coxa ye ll ow, mid and hind coxae (except apex) black; tip of hind femur blackish; fore tarsus with tip of tarsomere I a nd entire tarsomere 2 dark brown, tarsomere 3-4 black and s li ghtl y thickened, tarsomere 5 whi te; mid tarsus from tip of tarsomere I onward black; hind tarsus wholly black. Hai rs and bris tl es on legs black; fore coxa with 7 thi n or thick anterior and apical bristles; mid and hind coxae with I outer bris tl e; mid and hi nd femora with I preapical dorsal bri stle. Fo re tibia wi th I ad a nd 2 pd; mid ti bi a with 3 ad, 2 pd and I av; hind t ib ia with 4 ad, 4 pd a nd 3 av. Relative lengths of segments of legs Ll 3.2: 2.1: 1.3: 0.7: 0.4: 0.5; UI 4 .2 : 2 .3: 1.6 : 1.2: 0.7: 0.4; LIII 5.1: 1.4: 2.2: 1. 2: 0.7: 0.4. Wing hyaline,

Transcript of New and little known species of Dolichopodidae from China ...

Page 1: New and little known species of Dolichopodidae from China ...

BULLET! DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE E TOMOLOGI E. 70: 219-242. 2000 ENTOMOLOGIE. 70: 21 9,-242. 2000 BULLETIN VAN HET KONINK LIJ K BELGISCH INSTITUUT VOOR ATUU RWETENSCHAPI'EN

New and little known species of Dolichopodidae from China (VII): Diptera fro1n Etnei Mountain (2)

by DingY ANG and Toyohei SAIGUSA

Abstract

- 20 species of the family Dolichopodidae from Emei Mountain are described as new to science: !-lercos/omus (!-!.) curvarmalus, H. (!-!.) curvispinus. H. (H) incisus. 1-1. (!-!.) j indinganus, H (!-!.) longus. 1-1. (!-!.) 5piniwrsis, !-!. (H ) sub/angus, Dioslracus brevicercus, D. brevis. D. digiliformis, D. prolongalus. Amblypsilopus !ongisew . lvledelera curvala, Diaphorus lridenla/us. Rlwphium Jitrcalunl, Neurigonel!a !ongisew, N. pallipes. N. lri/itrcaw. Teuchoplwrus emeiensis, T. ven­/ralis.

Key wor·ds: Dolichopoclidae, new species, Emei Mountain, Sichuan

Introduction

II genera and 41 species of the family Dolichopodidae are known from Emei Mountain (YANG & SAIGUSA, 1999). Mainly based on the material collected by the second author, we add here 29 species to this region. Among them 20 species are described as new to science. The types of new species are mainly deposited in the China Agricultural University (CAU) except some para­types kept in theSAIGUSA collection at the Kyushu Uni­versity (SKU) and the '' Institut Royal des Sciences Nat­urelles de Belgique" (IRSN).

The fo llowing abbreviations are used: acr-acrostichal, ad-anterodorsal, apv-apicoventral, av-anteroventral , d­dorsal, dc-dorsocentral, h-humeral , ih-inner humeral, U-fore leg, LII-mid leg, LIII-hind leg, npl-notopleural, oc-ocellar, pa-postalar, pd-posterodorsal, ph-posthum­eral, psu-postsutural, pv-posteroventral, pvt-postvertical, sa-supraalar, su-sutural, v-ventral, vt-vertica l.

Subfamily Dolichopodinae

l. Plutlacrosoma briarea WEI et LJU

Phalacrosoma briarea WEI et LIU, 1996. J. Guizhou Agric. Coli. I 5(1 ): 38. Type locality: Anshun, Guizhou.

Specimens examined. I male, Sichuan: Emei Mountain, Maziba ( I, I 00- 1 ,300 m), 1998 . VIII. 16, T. SAIGUSA; 3 males, Sichuan: Emei Mounta in, Jingshui (700-750 m),

1998. VI II. 7 T. SAIGUSA; I male, Sichuan: Qingcheng Mountain, Yuechenghu (900 m), 1998. VIII. 22, T. SAI­GUSA (SKU).

Distribution. China (Sichuan, Guizhou).

2. Hercostomus (Hercostomus) curvispiuus sp. nov. (Figs. 1-2)

Diagnosis. Postocular bristles wholly black. First tlagel­lomere small and acute apically. Fore tarsomeres 3-4 black and slightly thickened, with thick dorsal hairs; tarsomere 5 whi te with white hairs.

Male. Body length 2 .8 mm, wing length 3.2 mm. Head metallic green with pale grey pollen; vertex,

frons and face with silvery white pollen; face nearly as wide as first tlagellomere. Hairs and bristles on head black; postocular bristles black, mid-ventral hairs pale. Antenna small and black; first tlagellomere as long as ~id)-4cute apically; _arista black, with basal segment 0. 17 t1mes as long as ap1cal segment, very short pubescent. Proboscis yellow with black hairs; palpus yellow, with black hairs and I black apical bristle.

Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 6 strong de; 5-6 irregularly paired acr short and hai r-like; scute llum with two pairs of bristles (outer pair very short, hair-like). Propleuron pale haired, with I black bristle on lower portion. Legs ye llow; fore coxa yellow, mid and hind coxae (except apex) black; tip of hind femur blackish; fore tarsus with tip of tarsomere I and entire tarsomere 2 dark brown, tarsomere 3-4 black and slightly th ickened, tarsomere 5 whi te; mid tarsus from tip of tarsomere I onward black; hind tarsus wholly black. Hai rs and bristles on legs black; fore coxa with 7 thi n or thick anterior and apical bristles; mid and hind coxae with I outer bristle; mid and hind femora with I preapical dorsal bristle. Fore tibia with I ad and 2 pd; mid tibia with 3 ad, 2 pd and I av; hind tibia with 4 ad, 4 pd and 3 av. Relative lengths of segments of legs Ll 3.2: 2.1: 1.3 : 0. 7: 0.4: 0.5; UI 4.2 : 2 .3: 1.6: 1.2: 0.7: 0.4; LIII 5.1: 1.4: 2.2: 1.2: 0 .7: 0.4. Wing hyaline,

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Figs. 1-2 - Hercostomus (Hercostomus) curvispinus, sp. nov., ma le. I. antenna (excluding scape), lateral view; 2. geni talia, lateral VIe W .

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Figs. 3-4 - !-!ercostomus (!-!ercostomw) incisus. sp. nov., male. 3. antenna (excluding scape). lateral view; 4. genital ia, lateral view.

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tinged with greyish; veins black, R4+5 and M distinctly convergent apically; CuAx ratio 0.5. Squama yellow with black hairs. Halter yellow, with dark brownish yellow stem.

Abdomen metallic green with pale grey pollen. Hairs and bristl es on abdomen black. Male genitalia (Fig. 2): Epandrium with long finger-like lateral lobe (bearing 2 apical bristl es); cercus rather small, with 2 apical denti­cl es; hypandrium irregularly furcate.

Female. Unknown.

Holotype male, paratype I male, Sichuan: Emei Moun­tain, Maziba (I, I 00-1 ,300 m), 1998. VIII. 16, T. SAIGUSA (CAU).

Distribution . China (Sichuan).

Etymology. The name refers to the curved spine on the hypandrium.

Remarks. The new species belongs to the hamatus­group. It is similar to dissectus Y A G et SAIGUSA from Sichuan, but may be di stinguished from the latter by the apically acute first tlagellomere. In dissectus , the first tlage llomere is obtuse apically (YANG & SAIGUSA, 1999).

3. Hercostomus (Hercostomus) henan us YANG

Hercostomus (Hercostomus) henanus Y A1 G, 1999. Bull. Inst. R. Sci. Nat. Bel g. Ent. 69: 210. Type locality: Henan.

Specimens examined. 2 males, Sichuan: Emei Mountain, Linggongli (I ,300-1,400 m), 1998. VIII. 17, T. SAIGUSA (SKU).

Distribution. China (Sichuan, Henan).

4. Hercostomus (Hercostomus) incisus sp. nov. (Figs. 3-4)

Diagnosis. Middle and lower postocular bristles pal e. First tlagellomere small and acute ap ically. Fore tarsome­res 3-4 bl ack, tarsomere 5 white. Mid tibia with I very long apv.

Male. Body length 3.8 mm, wing length 3.7 mm. Head metallic green with pale grey pollen; ve rtex,

fron s and face with silvery white pollen; face wider than first tlage ll omere. Hai rs and bri stles on head bl ack; mid and lower postocular bri stles (including ventral hairs) pal e. Antenna small and black; first nage ll omere as long as wide, acute ap ica lly; ari sta black, with basa l segment 0. 15 times as long a ap ica l segment, very short pubes­cent. Proboscis ye ll ow with black hairs; palpus yellow, with bl ack hairs and I black ap ica l bri stl e.

Thorax metallic green with pale grey pollen. Hairs and

bristles on thorax black; 6 sh·ong de; 6-8 irregularly paired acr short and hair-like; scutellum with two pairs of bri st les (outer pair very short, hair-like). Propleuron pale haired, with I black bristle on lower portion. Legs yellow; coxae yellow, mid coxa (except apex) blackish; tarsi from tip of tarsomere I onvvard dark brown, but fore tarsus with tarsomere 3-4 black, tarsomere 5 somewhat thickened and white. Hairs and bristles on legs black; fore coxa with 7 thin or thick anterior and apical bristles; mid and hind coxae with I outer bristle; mid and hind femora with I preapical dorsal bristle. Fore tibi a with I ad and 2 pel; mid tibia with 3 ad, 2 pel and I very long apv; hind tibia with 4 ad, 4 pel and 5 short av. Hind tarsomere I with 3 v. Relative lengths of segments of legs Ll 4.0: 2.1 : 1.4: 1.2: 0.7: 0.6; LII 5.4: 2.7: 1.7: 1.5: 0.9: 0.6; LIII 6.4: 1.7: 2.6: 1.6: 1.0: 0. 7. Wing hyaline, tinged with grey; veins black, R4+5 and M distinctly convergent apically; CuAx ratio 0.6. Squama yellow with black hairs. Halter yellow. Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black; venter with short pale hairs basa lly. Male genitalia (Fig. 4): Epandrium with lateral lobe nearly acute apically; cercus rather large, with dis­tinct apical incision; hypandrium irregularly furcate.

Female. Unknown.

Holotype male, Sichuan: Emei Mountain, Maziba (I , I 00-1,300 m), 1998. VIII. 16, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

Etymology. The name refers to the distinct apica l inci sion on the cercus.

Remarks. The new species is somewhat similar to curvis­pinus sp. nov. in the fore tarsus, but differs from the latter in the following points: middl e and lower postocular b1j trfs pale; mid tibia with I very long apv.

5. Hercostomus (Hercostomus) jindinganus sp. nov. (Figs. 5-6)

Diagnosis. Postocular bristl es wholly black. First tlage l­lomere obtuse ap ically. Male cercus with finger-like ap i­cal process.

Male. Body length 3.8 mm, wing length 4.6 mm. Head metallic green with pale grey poll en; face nar­

rower than first tlagellomere. Hairs and brist les on head black; postocular bristles black, mid-ventral hairs pale. Antenna small and black; first tlagellomere as long as wide, obtuse ap ica lly; ari sta black , with basa l segment 0.15 times as long as ap ica l segment, very short pubes­cent. Probosc is brown with black hai rs; palpus black , with black hairs and I black apica l bri stl e.

Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 6 strong de; 8-9 irregularly paired acr short and hair-like; sc utellum with two pairs ofbri ties

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Figs. 5-6 - Hercostomus (Hercostomus) jindinganus, sp. nov., male. 5. antenna, lateral view; 6. genital ia, lateral view.

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Figs. 7-8 - Hercostomus (Hercostomus) spinitarsis, sp. nov. , male. 7. antenna (excluding scape), latera l view; 8. genital ia, lateral view.

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(outer pair ve1y short, hair-like). Propleuron pale haired, with I black bristle on lower portion. Legs yellow; fore coxa yellow, mid and hind coxae (except apex) blackish to black; tarsi black, tarsomere I (except tip) dark brownish yellow. Hairs and bristles on legs black; fore coxa with 5-6 thin or thick anterior and apical bristles; mid and hind coxae with I outer bristle; mid and hind femora with I preapical dorsal bristle. Fore tibia with 2 ad and 2 pd; mid tibia with 2 ad, 2 pd and 2 av; hind tibia with 3 ad, 4 pd and 3 av. Hind tarsomere I with 3 v. Relative lengths of segments of legs LI 4.3 : 2.5: 1.35: 1.1 : 0.7: 0.7; Lll 6.3: 3.4: 2.1: 1.4: 0.8: 0.7; LIH 7.7: 2.4: 2.6: 1.6: 1.0: 0.75. Wing hyaline, slightly tinged with greyish; veins black, R4+5 and M strongly convergent apically; CuAx ratio 0.85. Squama yellow with black hairs. Halter yellow.

Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black; sternites 1-2 with some pale hairs. Male genitalia (Fig. 6): Epandrium with short lateral lobe (bearing I long apical bristle); cercus rather large, with finger-like apical process; hypandrium irre­gularly furcate .

Female. Unknown.

Holotype male, paratype I male, Sichuan: Emei Moun­tain, Jingding (2,900-3,030 m), 1998. VIII. 8, T. SAIGUSA (CAU & SKU).

Distribution. China (Sichuan).

Etymology. The species is named after the type locali ty Jingding.

Remarks . The new species is similar to digitatus YANG from Yunnan, but may be separated fi·om the latter by the arista with very short basal segment (0.15 times as long as apical segment). In digitatus, the arista is rather long (0.7 times as long as apical segment) (YA NG, 1997b ).

6. H ercostomus (Hercostomu!>) spinitarsis sp. nov. (Figs. 7-8)

Diagnosis. Postocular bristles wholly black. First flagel­lomere acute apically. Mid tarsus with spine-like ventral bristles.

Male. Body length 4.9 mm, wing length 4.6 mm. Head meta llic green with pale grey pollen; face slightly

narrower than first flagellomere. Hairs and bristles on head black; postocular bristles (including ventral hairs) wholly black. Antenna black; first fl agellomere 1.5 times longer than wide, acute apically; arista black, with basal segment 0.35 times as long as apical segment, short pubescent. Proboscis dark brownish ye llow with black hairs; palpus ye llow, with black hairs and I black apical bristle.

Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 6 strong de; 7-8 irregularly

paired acr short and hair-like; scutellum with two pai rs of bristles (outer pair very short, hair-like). Propleuron pale haired, with I black bristle on lower portion. Legs yellow; fore and hind coxae yellow, mid coxa (except apex) black; tarsi from tip of tarsomere I onward bro>vn­ish to brown. Hairs and bristles on legs black; fore coxa with 6 anterior and apical bristles, mid and hind coxae with I outer bristle; mid and hind femora with I preapical dorsal bristle. Fore tibia with I ad and 2 pd; mid tibia with 3-4 ad, 2-3 pd and 3 av; hind tibia with 4 ad, 4-5 pd and 5-6 av. Hind tarsomere I with 2 v at base. Mid tarsus nearly with n:vo rows of spine-like ventral bristles. Re­lative lengths of segments of legs LI 5.3: 2.7: 1.4: 1.1: 0.7: 0.5 ; Llf 6.7: 3.4: 1.7: 1.6: 1.0: 0.6; L III 7.8: 2.2: 2.9: 1.7: 1.0: 0.7. Wing hyaline, tinged with brown; veins black, R4+5 and M distinctly convergent apically. Squama yel­low with black hairs. Halter yellow. Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black. Male genitalia (Fig. 7): Epan­drium nearly acute apically; lateral lobe very slender, located near hypandrium; cercus wide apically with weak apical denticles; hypandrium irregularly furcate.

Female. Unknown.

!-!olotype male, Sichuan: Emei Mountain, Linggongli (1 ,300- 1,400 m), 1998. VIII. 17, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

Ety mology. The species is named after spine-like ventral bristles on the mid tarsus.

Remarks. The new species belongs to the digit([onnis­group. It is very similar to luoshanensis YANG et SAIGUSA from Henan. In luoshanensis, the mid tarsus has no spine-1/entral bristles (YANG & SAIGUSA, 2000).

7. H ercostomus (Hercostomus) weii Yang et Saigusa, nom. n.

1-Iercostomus Iii WEL, 1997. J. Guizhou Agric. Coli. 16(4): 35. Type locali ty: Yunnan. (preoccupied by 1-Ier­costomus Iii YANG, 1996. Ann. Soc. Ent. Fr. (N.S.) 32(4): 4 13)

Specimens examined. I male, Sichuan: Emei Mountain, Maziba ( I, I 00-1,300 m), 1998. Vfii . 16, T. SAIGUSA; 7 males, Sichuan: Emei Mountain, Linggongl i (I ,300-1,400 m), T. SAIGUSA (SKU).

Distribution . China (Sichuan, Yunnan).

Hercostomus (Hercostomus) longus-group

Diagnosis . Postocular bristles wholly black. Antenna wholly black. Scutellum with fi ne hairs on dorsal smface and apical margin. Mid and hind coxae wholly ye llow.

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Figs. 9- 11 - Hercos/omus (Hercoslomus) longus, sp. nov., male. 9. antenna (excluding scape), lateral view; I 0. hypandrium, ventral view; I I. geni talia, lateral view.

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Figs. 12- 14 - Hercostomus (Hercostomu.s) sub/angus, sp. nov. , male. 12. antenna, lateral view; 13. hypandri um, ventral view; 14. genitalia, lateral view.

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Mid and hind femora with some long ventra l hairs. Male gen italia : Epandrium elongated apically (much longer than wide), hypandrium partly hidden by epandrium; cercus long band-like, with some strong bristles.

The longus-group includes 4 following species: longus sp. nov., sub/angus, sp. nov, curvarmatus sp. nov. , ven­tralis Y A G et SAIG USA. Among them, H. longus and H. sub/angus are sister-species, which is evidently supported by the characters of the male genitalia.

8. Hercostomus (Hercostomus) longus sp. nov. (Figs. 9-11)

Diagnosis . Postocular bristles wholly black. First flagel­lomere slightly elongated, with angular apex. Mid and hind femora with long ventral hairs on basal half. Fore tibia with I very long apv .

Male. Body length 2.8-2 .9 mm, wing length 3.0-3. 1 mm. Head metallic green with pale grey pollen; face nar­

rower than first flagellomere. Hairs and bristles on head black; postocular bristles black, mid-ventral hairs mostly pale. Anten na black; first flagellomere ventra ll y with basal area brownish ye llow, 1.4 times longer than wide, angular apica lly; arista black, with basal segment 0.7 times as long as apical segment, very short pubescent. Proboscis brown to dark brown with black hairs; palpus black, with black hairs and I black ap ica l bristle.

Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 6 strong de; 6-7 irregularly paired acr short and hair-like; scute llum with two pairs of bristles (outer pair very short, hair-like), and with short hairs on dorsal surface and ap ical margin . Propleuron black and pale haired, with I black bristle on lower portion. Legs yellow; coxae ye ll ow, mid (except apex) blackish ; tarsi from tip of tarsomere I onward brownish to brown. Hairs and bristles on legs black; fore coxa with 7 thin or thick anterior and apical bristles; mid and hind coxae with I outer brist le, mid coxa with dense anterior hairs; mid and hind femora with I preapical dorsal bristle, and with long ventra l hairs on basal half. Fore tibia with I ad, 2 pd and I very long brown apv; n1id tibia with 3 ad, 2 pd and I av; hind tibia with 3 ad, 4 pd and 5 av. Hind tarsomere I with I vat base. Relative lengths of segments of legs Ll 2.7: 1.3: 0.7 : 0.6: 0.4: 0.35; LII 3.2: 1.7: 1.1: 0.9: 0.7: 0.4; LIII 4. 1: 1.3 : 1.7: 1.2: 0.75: 0.4. Wing hyaline, slight ly tinged with greyish; vei ns black, R4+5 and M convergent apically; CuAx ratio 0.5. Squama ye ll ow with black hairs. Halter yellow.

Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black. Male genita li a (Figs. 9-1 0) : Epandrium much elongated, ac ute apically, with short wide late ral lobe (bearing 2 lower apical bristles) ; cercus narrow, with 3 strong apica l bristles ; hypandrium irregularly furcate.

Female. Unknown.

Holotype male, paratypes 4 males, Sichuan: Emei Moun­tain, Maziba ( l , I 00-1 ,300 m), 1998. VI! f. 16, T. SAIGUSA; paratype I male, Sichuan: Qingcheng Mountain, Tian­rantuhua (900 m), 1998. VIII. 20, T. SAIG USA; l male, Sichuan: Qingcheng Mountain, Yuechenghu (900 m), 1998. VIII. 22 , T. SAIGUSA . (CAU , SKU & lRSN)

Distribution. China (Sichuan).

Etymology. The species is named after the mid and hind femora with long ventral hairs on basal half.

Remarks. The new species is similar to sub/angus sp. nov. in the mid coxa with dense anterior hairs and epandrium with slightly curved acute apex and apica lly separated latera l lobe bearing apical long hairs , but differs from the latter in the fore femur without dense ventral hairs and fore tibia with I ve1y long apv.

9. Hercostomus (Hercostomus) sublongus sp. nov. (Figs. 12- 14)

Diagnosis. Postocular bristles wholly black. First flagel­lomere slightl y elongated, with acute apex. Femora with long ventral hairs.

Male. Body length 2.6-2.7 mm, wing length 2.6-2.9 mm. Head meta llic green with pale grey pollen; vertex, frons and face with silvery wh ite pollen; face narrower than first flagellomere. Hairs and bristles on head black; posto­cul ar bristles black, mid-ventral hairs pale. Antenna black; first flagellomere 1.8 times longer than wide, acute apica lly; ari sta black, with basal segment 0. 7 times as long as apica l segment, very short pubescent. Proboscis dark brown with black hairs; palpus black, with black h~:s and I black apica l bristle.

/ Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 6 strong de; 6-8 iiTegularly paired acr short and hair-like; scutellum with two pairs of bristles (outer pair ve1y short, hai r-like), and with short hairs on cl01·sal surface and apical margin . Propleuron with black and pale hai rs on upper portion, pale hairs and l black bristle on lower portion. Legs yellow; coxae ye llow, mid coxa (except apex) tinged with blackish; tarsi from tip of tarsomere I onward brownish yellow. Hairs and bristles on legs black; fore coxa with 5 bristles on latera l and apical margin; mid and hind coxae with I outer bristle, mid coxa with dense anterior hairs; mid and hind femora with l preapical cl01·sal bristle. Fore femur with dense ventral hairs; mid femur with 2-3 long ventral hairs basally, hind femur with 4-6 long ventral hairs on basal half. Fore tibia with I ad and 2 pel; mid tibia with 3 ad, 2 pel and I av ; hind tibia with 2 ad, 3 pd and 4-5 av. Hind tarsomere I with l v at base. Relative lengths of segments of legs Ll 2.7: 1.3: 0.7: 0.4: 0.3: 0.35; Lll 3.5: 1.7: 1.2: 0.9: 0.7: 0.4; Llll 3.9: 1.4: 1.7: 1.1: 0.8: 0.4. Wing hyaline, slightly tinged with greyish; ve ins black, R4+5 and M slightly convergent apica lly; CuAx ratio 0.5 . Squama yellow with black hairs. Halter yellow.

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Figs. 15-17 - Hercostomus (Hercostomus) curvarmalus, sp. nov., male. 15. antenna, lateral view; 16. apical portion of hind tibia, lateral view; 17. genitalia, lateral view.

18 19

20

Figs. 18-20 - Dioslracus brevicercus, sp. nov., male. 18. palpus; 19. antenna, lateral view; 20. genitalia, lateral view.

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Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black. Male genitalia (Figs. 13-14) : Epandrium much elongated, acute apica lly; lateral lobe narrow apically, with 5 long apical bristles; cercus narrow apically, with some strong bri st les; hypandrium irregularly furcate.

Female. Unknown.

Holotype I male, Sichuan: Qingcheng Mountain, Yue­chenghu (900 m), 1998. Vlii . 22, T. SAIGUSA. Paratypes I male, Sichuan: Emei Mountain, Maziba ( I , I 00-1,300 m), 1998. VIII. 16, T. SAIGUSA; I male, Sichuan: Qingcheng Mot.ntain (900 m), 1998. VIII. 2 1, T. SAIGUSA . (CAU & SKU)

Distribution . China (Sichuan).

Etymology. The species is named after the simi larity with longus.

Remarks. The new species is a sister-species of longus sp. nov. , but may be separated from the latter by the fore femur with denser ventral hairs.

10. Hercostomus (Hercostomus) curvarmatus sp. nov. (Figs. 15-17)

Diagnosis. Postocular bristles wholly black. First flagel­lomere small and acute apica lly. Abdomen partly yellow basally. Hind tibia with T-shaped dorso-apical process.

Male. Body length 2.9 mm, wing length 3. 1 mm.

I ad, 2 pd and I av; hind tibia with 3 ad, 3 pd and 4-5 av, and with one T -shaped dorso-apical process. Relative lengths of segments of legs LI 2.7 : 1.5 : 0.7: 0.6: 0.3 : 0.35 ; Lll 3. 7: 1.8: 1.2: 0.9: 0.5: 0.4; UTI 4. 1: 1.4: 1.6: 1.0: 0.6: 0.4. Wing hyaline, tinged with greyish ; veins black, R4+5 and M weakly convergent apica lly; CuAx ratio 0.5. Squama yellow with black hairs. Halter ye llow.

Abdomen metallic green with pale grey pollen; tergites 1-4 tinged with yellow latera lly, sternites 1-4 ye llow. Hairs and bristles on abdomen black. Male genitalia (Fig. 17): Epandrium with indistinctly separated lateral lobe (bearing 3 strong apical bristles); cercus rather large, somewhat acute apica lly, with strong bristles; hypan­drium with nmTow curved lateral arm. Female. Unknown.

Holotype male, Sichuan: Emei Mountain, Maziba (I , I 00-1,300 m), 1998. VIII. 16, T. SAIGUSA (CAU).

Distribution . China (Sichuan).

Etymology. The species is named after the hypandrium with slightly curved lateral arm.

Remarks. The new species may be easi ly separated from other species of the longus-group by the mid tibia with T­shaped dorso-apical process.

Subfamily Hydrophorinae

11. Diostracus brevicercus sp. nov. (Figs. 18-20)

Head metallic green with pale grey pollen; vertex, frons Diagnosis. First flagellomere short, as long as wide; arista and face with silvery white pollen; face slightly narrower subap ical. Palpus go lden ye llow. 6 strong de. Legs chiefly than first flagellomere. Hairs and bristles on head black; black. Fore coxa with I black bristle on weak subapical postocular bristles (inc luding ventral hairs) wholly black. / tubercle. Antenna sma ll and black; first flage ll omere 1.1 times / longer than wide, acute apica ll y; arista black, with basal Male. Body length 4.5 mm, wing length 5.4 mm. segment 0.65 times as long as apical segment, very short Head metallic green with pale grey pollen; vertex pubescent. Proboscis blackish, with black hairs; palpus distinctly excavated. Eyes wide ly separated; face wi-black, with black hairs and I black apical bristle. dened downward. Hairs and bristles on head black; posto-

Thorax metallic green with pale grey pollen; meta- cul ar bristles black, lower postocular hairs (including pleuron brownish ye llow. Hairs and bristles on thorax ventra l hairs) mostly pale. Ocellar tubercle rather di stinct, black; 6 strong de; 6 irregularly paired acr short and hair- with one pair of rather long oc, without posterior hairs; I like; scutell um with two pairs of bristles (outer pair very long vt (longer than pvt). Antenna black; scape short, short, hair-like) , with short hairs on dorsal surface and without dorsal hairs; first flagellomere short subconical, apica l margin. Propleuron pale haired, with I long black as long as wide; arista subapical , one-segmented, black, bristle on lower portion . Legs ye llow; coxae yel low, mid very short pubescent. Proboscis black, with pale hairs; coxa with I blackish outer spot; fore tarsus from tip of palpus golden ye ll ow, with black hairs. tarsomere I onward brownish yellow; mid and hind tarsi Thorax metallic green with pale grey pollen; mesono-from tip of tarsomere I onward dark brown to black. tum with 2 indistinct brown longitudinal stripes between Hairs and bristles on legs black; fore coxa with 6 anterior de. Hairs and bristles on thorax black; 6 strong de; ac r and apical bristles; mid and hind coxae with I outer absent; I h, 1 ph, 2 npl , I sa, 1 pa, su and psu absent; bristle; mid and hind femora with I preapical dorsal scutellum with one pair of strong bristles and some very bri stl e. Mid femur with one row of 5 long black ventral short marginal hairs. Propleuron pale hairs , with I black hairs on basal half; hind femur with 3 long bl ack ventral bristle on lower portion. Legs black ; narrow apices of hairs basally. Fore tibia with I ad and 2 pd ; mid tibia with coxae and entire trochanters brownish yellow; narrow

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25

Figs. 2 1-25 - Diostracus brevis, sp. nov., male. 2 1. antenna, lateral view; 22. palpus; 23. hypandrium, ventral view; 24. genitalia, lateral view; 25. part of wing around m-cu.

Figs. 26-28 - Dio,1·1racus dig iti/armis, sp. nov. , male. 26. palpus; 27. antenna, lateral view; 28. gen ita li a, lateral view.

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bases of femora and tibiae, apices of femora ye llow. Hairs and bristl es on legs black; fore coxa with pale short hairs, and I bl ack bristle on weak subapical tubercle; femora with pale ventral hairs, but hind femur more or less with one row of very long weak ventral bristles. Fore tibia with 3 ad and I pd . Mid tibia with 4 ad and 3 pd, basa lly with some very long ventral bristles. Hind tibia with 5 ad, 3 pd and 5 weak av. Relati ve lengths of segments of legs LI 5.2: 2.2: 1.9: 1.3: 0.7: 0.7; LII 8.0: 3.8: 2.0: 1.3: 0.7: 0.7; LIII 8.5: 3.6: 2.6: 1.7: 1.0: 0.7. Wing hyaline, weakly tinged with greyish; ve ins black, R4+5 and M somewhat convergent; crossvein m-cu nearly straight. Squama brownish ye ll ow with mostly pale hairs. Halter ye llow.

Abdomen short and thick, as long as thorax, metallic green with pale grey pollen. Hairs on abdomen pale, but dorsum with mostly black hairs. Male genitalia (Fi g. 20): Epandrium rather wide, its surstylus long and strongly curved, acute apically; cercus short, Iobe-I ike; hypan­drium di stinctly curved, acute api cally.

Female. Body length 5.3 mm, wing length 6.6 mm. Similar to male, but palpus dark, mesonotum with 2 di st inct brown longitudinal stripes, hind femur and mi t tibia without long bristles.

Holotype male, paratype l fe male, Sichuan: Emei Moun­tain , Wuxiangang (700 m), 1998. VIII. 14, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

Etymology. The new spec ies is named afte r the short cercus.

Remarks. The new species is somewhat similar to neb­ulosus TAKAG I from Nepal and Tibet. But in nebulosus, the palpus is bl ack; 5 de (except 5th posterior de) are weak; the wing has one dark spot (TAKAG I, 1972; YANG, 1998).

12. Diostracus brevis sp. nov. (Figs. 2 1-25 )

Diagnosis. First fl age llomere rather short, as long as wide; ari sta nearly upper-apica l. Fore fe mur with one group of 7 posteri or bristles at base.

A-'faLe. Body length 5.6 mm, wing length 7.7 mm. Head metallic green with pale grey pollen; vertex

weakly excavated. Eyes widely separated; face widened downwa rd . Hairs and bri stl es on head black; postocular bri stl es bl ack, lower postocular hairs pale, ventral hairs rather long and pale. Ocell ar tubercle di stinct, with I pair of long oc, without posteri or hairs; l vt rather short; pvt strong, sli ghtly longer than vt. Antenna black, scape elongate, about 2 times longer than wide, wi th bl ack dorsa l hairs; firs t fl age ll omere rather short, as long as wide; ari sta subapical, one-segmented, bl ack, nea rl y bare.

Probosc is black, with pale hairs; palpus black vv ith grey brown poll en, with bl ack short hairs.

Thorax metallic green with pale grey pollen; mesono­tum somewhat dark , with 2 brownish bl ack longitudinal stripes between de, which extends backward near 5th posterior de. Hairs and bristles on thorax black; 6 de hair-like except 6th posterior de strong; acr absent; I h, I ph, 2 npl , I sa, I pa, su and psu absent; scutellum with 2 strong bristles. Propleuron pale haired, wi th I long pale hair-li ke bristle on lower portion. Legs entirely bl ack. Hairs and bri stl es on legs black; fore coxa with pale ye ll ow hairs (one group of basa l ciOI·sal hairs long and ye llow; one group of apical ciOI·sal bristles short, dense and bl ack); mid and hind coxae pale haired, except api cal ciOI·sa l hairs black. Fore femur basa lly with a group of 7 black long posterior bri stles; mid femur ventrally with 6 very long yellow hairs and I black bristle at base. Fore tibi a with 3 ad, 2 pel , a row of 4 long v (more or less at middl e); fore tarsomere I with one bomb of thick spine-li ke av and one row of long pv, fo re tarsomere 2 with one row of pv. Mid tibia with 3 ad and 2 pel. Hind tibia with 4 ad, 4 pa and 6-7 v; hind tarsomere I with 3 v. Relative lengths of segments of legs Ll 5.8: 3.0: 2.7: 1.3: 0.8: 1.0; LII 9.3: 5.4: 2. 1: 1.3: 0.8: 1.2; Llll 11 .6: 5.5: 3.5: 2.1: 1.1 : 1.2. Wing hyaline, slightly tinged with grey ish; ve ins black, R4+5 and M somewhat convergent; crossvein m-cu with a jet black nodule. Squama dark brown with pale hairs. Halter browni sh ye llow with black knob .

Abdomen short and thick, nearly as long as thorax; clark metallic green with pale grey poll en. Hairs on abdo­men pale and black. Male genitalia (F igs. 23 -24): Epan­drium rather wide apically, its surstylus with narrow apex curved and basa lly with I thin spine-like process; cercus slender with long hairs; hypandrium short, apica lly with obtuse late ral processes.

/ Female. Unknown.

HoLotype male, Sichuan: Emei Mountain, Linggongli ( I ,300- 1,400 m), 1998. VIII. 17, T. SAIGUSA (CAU). Distribution. China (Sichuan).

Etymology. The spec ies is named after the short first fl age llomere.

Remarks. The new species belongs to the unipunctatus­group. It is similar to saigusai TAKAG I from Taiwan. But in saigusai, the fo re femur has only 1-2 poste rior bri stles at base and fo re tibia has 3 long pv at middle (TAKAG I, 1968).

13. Diostracus digit(formis sp. nov. (F igs. 26-28)

Diagnosis . vt ab ·ent. First fl age ll omere elongate, 1.6 times longer than wide; arista upper ap ica l. Legs wholly bl ack. Empodium and pul villi indistinct.

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31

Figs. 29-33 - Dioslracus prolongatus, sp. nov., male. 29. pa lpus; 30. antenna, lateral view; 31 . part of wing around m-cu; 32. hypandrium, ventral view; 33. geni talia, latera l view.

Male. Body length 6.3 mm, wing length 6.6 mm. Head metallic green with grey brown pollen, occiput

pale grey pollinose except upper portion; vertex weakly excavated. Eyes widely separated; face widened down­ward. Hairs and bristles on head black; 5 strong posto­cular bristles (2 upper ones black, 3 lower ones yellow), lower postocular hairs (including ventral hairs) pale. Ocellar tubercle distinct, with one pair of strong oc, without posterior hairs; vt absent, pvt slightly shorter than oc. Antenna black; scape short, without dorsal hairs; fi rst flagellomere elongate, 1.6 times longer than wide; arista upper apical, one-segmented, black, nearly bare. Probos­cis black, with pale hairs; palpus black, with black hai rs. Thorax metallic green with pale grey pollen; mesonotum with 2 brown longitudinal stripes between de (which reach 5th posterior de). Hairs and bristles on thorax black; 6 de hair-li ke except 6th posterior de strong; acr absent; I h, I weak ph nearer to transverse suture than to h, 2 npl, I sa, I pa, su and psu absent; scutellum with one pair of strong bristles and 3 very short hairs . Propleuron with I pale hair on upper portion, I pale hair and I pale long weak bristle on lower portion. Legs wholly black; empo­dium and pulvill i reduced and indistinct. Hairs and bris­tles on legs black; fo re coxa with pale hairs, and with brownish yellow bristles at anterior apical margin. Fe­mora with some pale ventra l hairs, but hind femur with pale and a little long ventral hairs (nearly as long as width

offemur). Fore tibia with 4 ad, 2 pd and 8 pv. Mid tibia with 3 ad, 2 pd, 4 av, 5 pv and 4 v. Hind tibia with 4 ad, 2 pa, 6 av and 5-6 pv. Relative lengths of segments oflegs Ll 6.4: 1.6: 2.5: 1.4: 0.8: 1.2; LII 8.9: 3.8: 1.7: 1.0: 0.7: 1.0; LIII 9.8: 4.4: 2.7: 1.5: 0.8: 1.0. Wing hyaline, indis-

)mctly tinged with greyish; veins black, R4+5 and M / somewhat convergent; crossvein m-cu nearly straight.

Squama dark brown with pale hairs. Halter yellow. Abdomen longer than thorax, metallic green with pale

grey pollen. Hairs on abdomen pale, but dorsum with mostly black hairs. Male genitalia (Fig. 28): Epandrium rather wide, its surstylus short and broad; cercus long finger-like; hypandrium rather wide in lateral view. Female. Unknown.

Holotype male, Sichuan: Emei Mountain, Jingshui (700-750 m), 1998. VIII. 15, T. SAIGUSA (CAU).

Distribution. China (S ichuan).

Etymology. The species is named after the long finger­like cercus.

Remarks. The new species belong to the fenestra/us­group vvith 3 known species (SAIGUSA, 1984). It may be easily separated from them by the wing without any accessory cellulae.

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14. Diostracus prolongatus sp. nov. (Figs. 29-33)

Diagnosis. First tlagellomere elongate, 1.6 times longer than wide; arista subapical. Fore femur with one group of 4 posterior bristles at base.

Male. Body length 6.3 mm, wing length 7.7 mm. Head metallic green with pale grey pollen; vertex weakly

excavated. Eyes widely separated; face widened down­ward. Hairs and bristles on head black; middle and lower postocular bristles pale yellow, ventral hairs rather long and pale. Ocellar tubercle with one pair of long oc, without posterior hairs; I short vt, pvt longer than vt. Antenna black; scape elongate, about 2 times longer than wide, with black dorsal hairs; first flagellomere elongate, 1.6 times longer than wide; arista subapical, one-segmented, black, nearly bare. Proboscis black, with pale hairs; palpus black with grey brown pollen, with black sh01i hairs.

Thorax metallic green with pale grey pollen; mesono­tum somewhat dark, with 2 brownish black longitudinal stripes between de, which reaches 5th posterior de. Hairs and bristles on thorax black; 6 de hair-like except 6th posterior de strong; acr absent; I h, I ph, 2 npl , I sa, I pa, su and psu absent; scute llum with one pair of strong bristles and 2 very short hairs. Propleuron pale hairs, with I pale hair-like bristle on lower portion. Legs entirely black. Hairs and bristles on legs black. Fore coxa pale haired, with long yellow hairs at narrow base and 6-7 black bristles near anterior distal margin. Femora with some pale ventral hairs; mid femur with distinct yel low ventral hairs including more or less one row of 7-8 very long hair- li ke pv on basal 1/3. Fore femur basally with a group of 4 black long posterior bristles. Fore tibia with 4 ad, 2 pd, a row of I 0 pv; fore tarsomere I with one comb of short thick spine-like av and one row of long pv, fore tarsomere 2 with one row of pv on whole length and one sh01i row of 7 short spine-li ke av at base. Mid tibia with 3 ad and 2 pd. Hind tibia with 5 ad, 4 pa, 4 av and 3 pv. Relative lengths of segments of legs LI 6.0: 3.0: 2.6: 1.5: 1.0: 1.1; LII 9.0: 5.0: 2. 1: 1.4: 0.85: 1.2; LilT 11.8: 5.1: 3.6: 2.2: 1.1: 1.2. Wing hyaline, weak ly tinged with greyish ; veins black, R4+5 and M somewhat con­vergent; crossvein m-cu with a jet black nodule. Squama dark brown with pale hairs. Halter brown with black knob.

Abdomen thick, slightly longer than thorax; dark me­tallic green with pale grey pollen. Hairs on abdomen mostly pale; tergites 4-5 with long go lden yellow hairs laterally; cercus with long go lden yellow hairs. Male genitalia (Figs. 32-33): Epandrium rather wide apically, its surstylus thick and nearly straight with I thin spine­like process; cercus slender with long hairs; hypandrium short, apically with acute lateral processes.

Female. Unknown.

1-!o/otype male, Sichuan: Emei Mountain, Jingding (2,900-3,030 m), 1998. VIII. 8, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

Etymology. The new species is named after the elongate first tlagellomere.

Remarks. The new species belongs to the unipunctatus­group. It is simi lar to longicornis SAIGUSA from Nepal in the shape of the first tlagellomere. In longicornis, the fore femur has 1-2 posterior bristles at base, the 3rd abdominal sternum has long dense golden yellow hairs (SAIGUSA, 1984).

Subfamily Sciapodinae

15. Condylostylus luteicoxa P ARENT

Condylosty lus luteicoxa PARENT, 1929. Ann. Soc. Sci. Brux. 49: 225. Type locality: Assam, India.

Specimens examined. I male I female , Sichuan: Emei Mountain, Maziba (I, 1-1 ,300 m), 1998. VIII. 16, T. SAIGUSA; 1 male, Sichuan: Emei Mountain, Jingshui (700-750 m), 1998. VIII . 7 T. SAIGUSA; I female , Si­chuan: Emei Mountain, Linggongli ( 1,300-1,400 m), 1998 . VIII. 17, T. SAIGUSA; I fema le, Sichuan: Qing­cheng Mountain (850-I ,000 m), 1998. VIII. 21, T. SAI­GUSA (SKU).

Distribution. China (Sichuan, Guizhou, Yunna, Zhejiang, Fujian, Taiwan, Henan); Japan, India.

16. Amblypsilopus longiseta sp. nov. (Figs. 34-35)

Diagnosis . Frons with 1 very long black anterior bristle / en postero-lateral slope. Palpus brownish yel low.

Male. Body length 4.2 mm, wing length 3.7 mm. Head brilliant metallic green with pale grey pollen.

Vertex distinctly excavated; eyes widely separated; cly­peus separated from eye margin. Hairs and bristles on head black; middle and lower postocular bristles (includ­ing ventral hairs) pale. Ocellar tubercle distinct, with one pair of long oc and 2 very short posterior hairs ; frons with 1 very long anterior bristle and 2 long posterior bristles (inner bristle slightly longer than outer bristle) on pos­tero-lateral slope. Antenna black; first tlagellomere nearly triangular, as long as wide; arista apical , black, slightly longer than width of head. Proboscis brownish yel low, with pale hairs; palpus brownish yellow, with pale hairs, I long black subapical bristle and 1 short black apical bristle.

Thorax brilliant metallic green with pale grey pollen. Hairs and bristles on thorax bl ack; 3-4 short anterior de hair-like and 2 strong posterior de; 2 long irregularly paired acr; h absent, 1 ph , 2 long npl, I short su, 2 sa (anterior sa short), 1 pa. Scutellum on ly with one pair of

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Figs. 34-35 - Amblypsi/opus /ong iseta, sp. nov., male. 34. hypandrium, ventral view; 35. genitalia, lateral view.

36 ~ / 37

38

39

Figs. 36-39 - Medetera curvata, sp. nov. , male. 36. hypandrium, ventral view; 3 7. ap ical portion of aedeagus, lateral view; 38. genita lia, latera l view; 39. cercus, latera l view.

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bristles (without latera l bristles). Legs ye llow; fore coxa ye ll ow, mid and hind coxae black; tarsi from tip of tarsomere I onward brown to dark brown. Hairs and bristles on legs black; coxae with pale hairs and bristles; femora with pale ventra l hairs . Mid tibia with I very short ad and l very short pd at base; hind tibia with 7-8 very short pd and 8-9 very short av. Relative lengths of seg­ments of legs LI 3.7: 3.2: 1.2: 0.8 : 0.5 : 0.4; Lll 4.7: 4.4: 1.2: 1.0: 0.6: 0.35 ; Llll 6.9: 3.0: 1.4: 1.0: 0.6: 0.35 . Wing hyaline; veins black; M I with short base distinct. Squama brownish ye llow with dark brown margin, with black hairs. Halter ye llow with dark brown stem.

Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black; tergite I with pale hairs latera lly. Male gen italia (Figs. 34-35): Epandrium longer than wide, surstylus short and more or less tapering toward to tip (bearing 3 bristles), latera l lobe small with thin apex (bearing 2 bristles) ; cercus nan·ow and slightly curved.

Female. Unknown.

Ho/otype male, Sichuan : Emei Mountain, Jingshui (650 m), 1998. Vlll. I 0, T. SAIGUSA (CAU).

Distribution . China (S ichuan).

Etymology. The species is named afte r the frons with very long bristle.

Remarks. The new species is simil ar to sichuanensis YANG, but differs from the latter in the frons with I very long bri stle and pal pus brownish yellow. In sichuanensis, the frons has I distinct hair, the pal pus is black (YANG, 1997a).

Subfamily Medeteriuae

17. Medetera curvata sp. nov. (Figs. 36-39)

Diagnosis. Postocular bristles who ll y black. Propleuron with 3 black brist les. Antenna brownish ye llow or ye ll ow with black first flagellomere.

Male. Body length 4.3 mm, wing length 4.7 mm. Head metallic green with pale grey pollen; upper occi­

put concave. Eyes distinctly separated; face narrower than frons. Hairs and bristles on head black; postocular br ist les who lly black. Oce ll ar tuberc le weak, with one pair of very long oc and 2 very short posterior hairs; I long vt. Antenna! scape and pedicel brownish yellow; first flagellomere black, somewhat ova l, nearly as long as wide, with short hairs ; arista nearly ap ica l, very long, black, with very short pubescence. Probosc is large and thick, black, with sparse black hairs; palpus metallic green with brownish hairs and I black subapical brist le.

Thorax meta lli c green with pale grey po llen; mesono-

tum and scutellum with grey brown pollen. Hairs and bristles on thorax black; 5 de; 6-7 irregularly paired acr short and hair-like; I h and 2 short pale hairs, I ph, 2 npl , 2 sa, I pa; scutellum with two pairs of bristles. Propleuron with 3 black bristles. Legs black (hind tarsus broken); femo ra with yellow narrow apices; tibiae with brownish ye llow extreme bases; f01'e tibia and tarsus blackish. Hairs and bristles on legs black; hind coxa with I outer bristle at middle. Hind femur with one row of 6 dorsal bristles basally and 3-4 av apically; mid tibia with I ad and I pd basally. Relative lengths of segments of legs LI 4.1: 2.2: 1.3 : 0.8: 0.5: 0.6; Lll 5.4: 2.5: 1.6: 1.2: 0.6: 0.5 ; Llll6.7:? :?:?:?:?. Wing nearly hyaline; veins dark brown, R4+5 and M strongly convergent apica lly; CuAx ratio 0.5. Squama ye llow with brownish margin, and with black hairs. Halter yellow.

Abdomen metallic green with grey brown pollen. Hairs and bristles on abdomen mostly black; tergite I with longer hairs and bristles. Ma le genitalia (Figs. 36-39): Epandri um distinctly elongated, with 2 finger-like lateral processes (bearing I long bristle) , its surstylus rather thick with 2 curved apical projections; cercus rather narrow apica lly, with a obtuse ap ica l process; hypan­drium long and nearly acute apically; aedeagus with short thi n curved tip.

Female. Body length 3.7 mm, wi ng length 4.2 mm. Similar to male, but antenna! scpae and pedicel ye llow, femora with a little wide apex ye llow, tibiae brownish ye llow.

Ho/otype male, paratype I female , Sichuan: Emei Moun­tain, Leiclongping (2 ,400 m), 1998. VIJf. 13, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

IJ~ology. The species is named after the surstylus with curved apical processes.

Remarks. The new species is similar to neo::wngensis YANG et SAIGUSA from Henan. It may be easi ly separated from the latter by having the propleuron with 3 bristles and sursty lus with narrow apical processes distinctly curved.

18. Paramedetera medialis YANG et SAIGUSA

Paramedetera medialis YANG et SAIGUSA, 2000. In: In­sects of the Funiu Mountains region (2) . Type locality: Henan.

Specim.ens examined. I male, Sichuan: Emei Mountain, Wuxiangang (700 m), 1998. Vlll. 14, Ding YANG ; 3 males I female, Sichuan: Emei Mounta in, Jingshui (650 m), 1998. Vll I. I 0, Ding YANG (CAU).

Distribution. China (Sichuan, Henan).

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234 by DingY ANG and Toyohei SAIGUSA

Figs. 40-4 1 - Diaphorus tridentatus, sp. nov., male. 40. antenna, lateral view; 41. genitalia, lateral view.

42

\ \

\

43

_> --.... ..," ..... .>."' ' ..... ,

' I ' ' I ' ',\ \

40

41

Figs. 42-43 - Rhaphiumji./1 -ccttum. sp. nov. , male. 42. antenna (exc luding scape), lateral view; 43. genitalia, latera l view.

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Dolichopodidae from China (VTII) : Emei moutain I I 235

Subfamily Diaphoriuae

19. Diaplwrus nigricans MEIGEN

Diaphorus nigricans MEIG EN, 1824. Syst. Beschr. 4: 33. Type locali ty: not given.

Specimen examined. I male, Sichuan: Emei Mountain, Linggongli ( 1,300-1,400 m), 1998. VHl . 17, T. SAIGUSA (SKU).

Distribution. China (Sichuan, Henan); Palaearctic, Nearctic and Neotropical regions.

20. Diaphorus lichwardti P ARENT

. Diaphorus lichwardti PARENT, 1925. Ann. Soc. Sci. Brux. 44: 265. Type locality: Kou-Ling, Jiangx i.

Specimen examined. I male, Sichuan: Emei Mountain, Ma­ziba (1,100-1 ,3 00 m), 1998. VIII. 16, T. SAIGUSA (SKU).

Distribution. China (Sichuan, Jiangx i) .

21. Diaphorus trideutatus sp. nov. (Figs. 40-41)

Diagnosis. Postocular bristl es wholly black. Hind femur apically wi th long thin ventral hairs. Lateral lobe on epandrium with 3 marginal denticles.

Male. Body length 3.6-4.0 mm, wing length 3.3-3.9 mm. Head metallic green with grey brown pollen. Eyes con­

tiguous on frons for a long distance (about 1/2 of frons) , with dorsa-anterior facets enlarged. Hairs and bristl es on head black; postocular bristles wholly black. Ocellar tu­bercle distinct, with one pair of long oc and 2 very short posterior hairs; I short vt . Antenna black; first flagellomere 1.6 times wider than long; arista subapical, black, very short pubescent. Proboscis black, with black hairs; palpus yellow, with black hairs and I bl ack apical bristle.

Thorax meta llic green with pale grey poll en. Hairs and brist les on thorax black; 5 strong de; 9-1 0 irregularly paired acr short and hair-like; I h and 2 very short hairs, I ph , I ih, I su, 2 npl , 2 sa, I pa; scutellum wi th two pairs of bristles (outer pair very short, hair-like). Prop leuron with 2 black hairs and I long black bri stle on lower portion . Legs black; fore and mid femora with ap ical 1/5 yellow, hind femur with extreme tip brownish yellow; tibi ae yellow except apex of hind tibi a black ish; fore and mid tarsi yellow, except fore tarsomeres 3-4 black ish and tarsomere 5 black, mid tarsus from tip of tarsomere I onward brown to dark brown. Hai rs and bristles on legs black ; fore coxa with 6 anterior and ap ica l bristles; mid coxa with 2 outer brist les; hind coxa wi th I outer bri st le at base. Fore fem ur with one row of weak av basally and one row of weak pv; hind femur api ca ll y with 3 thick av and

some long ventral hairs. Fore tibia with I ad and 2 pd; mid tibia with I ad, 2 pd and I av; hind tibia with I weak ad, 5 pd and 5 short thin av. Fore tarsus without claws, with elongated pulvilli which are a little longer than tarsomere 5; mid and hind tars i with 2 small claws and 2 small pulvilli . Relative lengths of segments of legs LI 3.8: 2.2: 1.2: 0.8: 0.6: 0.5; LII 4.5: 2.5: 1.4: 1.0: 0.6: 0.4; LIII 5.8: 1.6: 1.7: 1.25: 0.7: 0.45 . Wing hyaline; veins dark brown, R4+5 and M parallel apically; CuAx ratio 0.5. Squama yellow with black hairs. Halter yellow with brown base.

Abdomen metallic green with pale grey pollen. Hairs and bristles on abdomen black; sternite 8 with 4 strong bristles. Male genitalia (Fig. 41): Epandrium with lateral lobe rather large (bearing 3 marginal denticles), its sur­stylus long and distinctly narrow apically; cercus long with narrow base.

Female. Unknown .

Holotype male, paratype I male, Sichuan: Emei Moun­tain, Jingshui (700-750 m), 1998. VIli. 15, T. SAIGUSA. (CAU & SKU)

Distribution. China (Sichuan).

Etymology. The species is named after the lateral lobe on epandrium with 3 marginal denticles.

Remarks. The new species is very similar to menglunen­sis YANG et GROOTAERT from Yunnan. It may be sepa­rated from the latter by having the cercus rather wide with narrow base and lateral lobe on epandrium with three distinct marginal denticles. In menglunensis, the cercus is uniformly thick, the lateral lobe on epandrium has I weak marginal denticle (YANG & GROOTAERT, 1999).

Subfamily Rl!aphiinae

22. Raphium dispar COQUJLLET

Raphium dispar COQU ILL ET, 1898. Proc . U. S. Nat. Mus. 21: 3 19. Type locality : Japan.

Specimens examined. I male, Sichuan: Emei Mountain, Linggongli ( 1,300-1 ,400 m), 1998. VIII. 17, T. SAIGUSA; I male, Sichuan: Qingcheng Mountain, Yuechenghu (900 m), 1998. Vlii . 22 , T. SAIGUSA (SKU).

Distribution. China (S ichuan, Guizhou, Zhejiang, Tai­wan); Russ ia (Far East), Japan.

23. Rhaphium jitrcatum sp. nov. (F igs. 42-43)

Diagnosis. First flagellomere long ( 4.5 times longer than wide); arista rather short (0.5 times as long as fi rst fla-

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236 by Ding YANG and Toyohei SAIGUSA

gellomere). acr absent. All coxae yellow. Surstylus fur­cate apically.

Male. Body length 1.5 mm, wing length 2.0 mm. Head metallic green with pale grey pollen. Frons rather

wide; face distinctly narrower than frons, narrowing downward; eyes somewhat contiguous above oral mar­gin. Hairs and bristles on head black; middle and lower postocular bristles (including ventral hairs) pale. Ocellar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna black; fi rst fl agellomere long ( 4.5 times longer than wide); ari sta apical, rather short (0.5 times as long as first fl agellomere). Proboscis brown­ish, more or less with pale hairs; palpus yellow, with pale hairs and 2 short black bristles.

Thorax metallic green with pale grey pollen. Hairs and bristles on thorax black; 5 de strong except 4th posterior de shOJi and weak; 2-3 acr present only before first ante­rior de; I h and I ve1y sho1i hair, I ph, I ih, I weak su, 2 npl, 2 sa (anterior bristle short), I pa; scutellum with two pairs of bristles (outer pair ve1y shOJi , hair-like). Pro­pleuron with I pale hair on upper pOJiion and I weak pale bristle on lower pOJiion. Legs yellow; all coxae yellow; tarsomeres 2-5 brownish yellow. Hairs and bristles on legs black; fore coxa with 4 bristles on apical margin; mid coxa with I outer bristle anteriorly; hind coxa with I outer bristle at middle. Mid femur with I preapical anterior bristle; hind femur apically with 2 av. Mid tibia with 2 ad, 2 pd and I av;

'

''

hind tibia with 2 ad, 3 pd and I av. Relative lengths of segments oflegs U 1.7: 0.9: 0.65: 0.6: 0.3: 0.25; LII 2.6: 1.2: 0.7: 0.6: 0.4: 0.25; LIJI 3.0: 0.75: 0.8: 0.7: 0.4: 0.3. Wing hyaline; veins brown, R4+5 and M convergent apically; CuAx ratio 0.4. Squama yellow with pale and blackish hairs. Halter yellow.

Abdomen metallic green with grey pollen; cercus brownish. Hairs and bristles on abdomen black. Male geni talia (Fig. 43): Epandrium somevvhat rounded, its surstylus bent and furcate apically; cercus very long and thin apically.

Female. Unknown.

Holotype male, Sichuan: Emei Mountain, Maziba (I, I 00-1,300 m), 1998. VIII. 16, T. SAIGUSA (CAU).

Distribution. China (Sichuan).

Etymology . The species is named after the furcate surstylus.

Remarks. The new species is somewhat simi lar to apici­nigrum YANG et SAIGUSA from Sichuan, but may be distinguished from the latter in the following points: palpus yellow, hind femur not dark apically, cercus with­out lateral lobe; in apicinigrum, palpus black, hind femur with black apex; cercus with short lateral lobe (YANG & SAIGUSA, 1999).

- -=.:=----- ::--

' ' ' ' , _

... ..... - --

' ' ' \ \ I I I I I I

I I

I I

44

Fig. 44 - Neurigonella longiseta, sp. nov., male genita lia, lateral vievv.

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Dolichopodidae from China (VIII): Emei moutain 23 7

Subfamily Peloropeodinae

24. Neurigonella longiseta sp. nov. (Fig. 44)

Diagnosis. All coxae yellow. Halter pale ye llow. Cercus with basa l tubercle bearing very long brist les.

Male. Body length 1.8 mm, wing length 3.1 mm. Head metallic green with pale grey pollen. Eyes di s­

tinctly separated. Frons rather wide; face distinctly nar­rower than frons , narrowing downward. Hairs and bri st les on head black; postocular bri stl es (including ventral hairs) bl ack. Ocellar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna black; first fl age llomere nearly trapezo id, nearly as long as wide, incised api ca lly; ari sta black with very short basa l seg­ment. Probosc is black, with bl ack hairs; palpus blac k, wi th bl ack hairs and I bl ack bristle.

Thorax metallic green with pale grey pollen; mesono­tum with brown pollen. Hairs and bristles on thorax black; 5 de strong, 4-5 irregul arly pai red acr short and hair-like; 1 h and I short hair, I ph, I ih, 1 su, 2 npl , 2 sa (anterior bristle short), I pa; scutellum with two pairs of bri stles (outer pair very short, hair-like). Propleuron with 2 short pale hairs on upper portion, I short pale hair and I long bl ack bristle on lower portion. Legs yellow; all coxae yellow; tarsomere 5 dark brown. Hairs and bristles on legs black; fore coxa with 5 bristles; mid coxa with I outer bristle anteriorl y; hind coxa with I outer bristle at middle. Mid and hind femora with I preapical bristle. Mid tibia with 2 ad and 2 pd ; hind tibia with I ad and I pd . Relative lengths of segments of legs U 1.7: 0.8: 0.7: 0.4: 0.3: 0.3 ; LII 2.6: 1.35 : 0.8: 0.7: 0.4: 0.3 ; Llll 2.8: 0.7: 1.2: 0.8: 0.4: 0.3. Wing hya line; ve ins dark brown, R4+5 and M slightly convergent apically; CuAx ratio 0.45 . Squama yellow with dark brown margin, with black hairs. Halter pale yellow.

Abdomen metallic green with pale grey poll en. Hairs and bristl es on abdomen black; venter with pale hairs . Male genitalia (Fig. 44): Epandrium with 2 lateral bris­tles, of which I brist le is very long and located on the tubercl e; cercus with basa l tubercle with short hairs and 2 very long bri stles.

Female. Unknown.

Holotype male, Sichuan: Emei Mountain, Linggongli ( I ,300-1 ,400 m), 1998. V!IT. 17, T. SAJGUSA (CAU).

Distribution. China (S ichuan).

Etymology. The spec ies is named after the cercus with very long bri stles.

Remarks. Only I spec ies of the genus Neurigonella Ro­BINSON was known to occur in the Palaearct ic region: tatjanae NEG ROBOV, 1984 from Far East of Russia (N E­ROBOV, 199 1 ). YANG, YANG & Li ( 1998) reported the first Chinese spec ies henanensis fro m Henan. Here we de­scribe 3 new species from Emei Mountain, Sichuan.

The new species is similar to pa/lipes sp. nov., but may separated from the latter by having the halter pale yellow and fore tarsomere 5 dark brown.

25. Neurigonella pallipes sp. nov. (Fig.45)

Diagnosis . All coxae brownish ye llow. Fore tarsomere 5 white. Halter bl ack with ye llow base.

Male. Body length 2.3 mm, wing length 2.3 mm. Head metallic green with grey brown poll en. Eyes

distinctly separated. Frons rather wide; face distinctly narrower than frons , narrowing downward. Hairs and bristles on head bl ack; postocular bristles (including ven­tral hairs) black. Ocellar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna black ; first fl age llomere nearly trapezoid, 0.9 times longer than wide, incised apically; ari sta black, with very short basa l segment. Probosc is black, with black hairs; palpus black, with black hairs and I black bristle.

Thorax metallic green with grey brown pollen. Hairs and bristles on thorax black; 5 de stmng, 6 irregularly paired acr sh01t and hair-like; I h and I sh01t hair, I ph, I ih, I su, 2 npl , 2 sa (anterior bristle short), I pa; scutellum with two pairs of bristles (outer pair ve1y sh01t, hair-like). Prop lemon with I sh01t pale hair and I long black bristle on lower p01tion. Legs brownish yellow; tarsi from tip of tarsomere I onward brown to dark brown; fore tarsomere 5 white. Hairs and bristles on legs black; fore coxa with 7 bristles; mid coxa with I outer bristle anteriorly; hind coxa with I outer bristle at middle. Mid and hind femora with I preapical anterior bristl e. Mid tibia with 2 ad and 2 pd; hind tibia with I ad and 2 pd. Relative lengths of segments of legs Ll 2.2: 1.3: 0.8: 0.8 : 0.7: 0.35; LII 3.2: 1.8: 1.1: 0.9: 0.7: 0.35; LI!I 3.7: 0.7 : 1.6: 1.1 : 0.7: 0.35. Wing hyaline, weakly tinged with greyish; veiysdark brown, R4+5 and M slightly convergent apically; CoAx ratio 0.4. Squama yellow with dark brown margin, with black hai rs. Halter black with yellow base.

Abdomen metallic green with grey brown pollen. Hairs and bri stles on abdomen black. Male genitalia (Fig. 45): Epandrium with 2 lateral bri stl es, of which I bristle is very long and located on weak tubercle; cercus with one group of short hairs basally.

Female. Unknown.

J-lolotype male, Sichuan: Emei Mountain, Linggongli ( 1,300-1,400 m), 1998 . vm. 17, T. SAJGUSA (CAU).

Distribution. China (S ichuan).

Ety nw logy . The species is named after the fore tarsomere 5 white.

Remarks. The new spec ies is similar to longiseta sp. nov, but may be separated from the latter by hav ing the fore tarsomere 5 white and halter black with yellow base.

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238 by Ding YANG and Toyohei SAIGUSA

... ... ....

Fig. 45 - Neurigonel!a pal!ipes, sp. nov., male genitalia, lateral view.

46

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' J ) ' ....

' ' /

;~ ,_-; ".-_~:.:,-~: ~ '--

.... ' ' ' '

48

' ' ' \ \

45

\ \ I I I I

, I

I

I I

Figs. 46-48 - Neurigonella trijitrcata, sp. nov., ma le. 46. wing; 47. antenna (exc luding scape and pedicel), lateral view; 48. genita lia, latera l view.

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Dolichopodidae from China (VIII): Emei moutain, , 239

26. Neurigonella trifurcata sp. nov. (Figs. 46-48)

Diagnosis. Mid and hind coxae (except apex) blackish. R2+ 3 and R4+5 with sh01t bases distinctly thickened. Halter dark brown. Cercus with short or long hairs ba­sally . Aedeagus trifurcate apically.

Male. Body length 2.4 mm, wing length 2.2 mm. Head metallic green with grey pollen. Eyes distinctly

separated. Frons rather wide; face distinctly narrower than frons, narrowing downward. Hairs and bristles on head black; postocular bristl es (including ventral hairs) black. Ocellar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna black; first fl agello­mere nearly trapezoid, nearly as long as w ide, incised apically; ari sta black, with very short basal segment. Proboscis brown, w ith black hairs; palpus black, with black hairs and I black bristle.

Thorax metallic green with grey pollen. Hairs and bristles on thorax black; 5 de strong, 7 iiTegularly paired acr short and hair-like; I hand 1 short hair, I ph, 1 ih, I su, 2 npl , 2 sa, I pa; scute llum with two pairs of bristles (outer pair very short, ha ir-like). Propleuron w ith I short black hair on upper portion, 3 short black hairs and I long black bristl e on lower portion. Legs yellow; fore coxa yel low, mid and hind coxae (except apex) blackish. Hairs and bristles on legs black; fore coxa with 7 bristles; mid coxa w ith I outer bristle anteriorly; hind coxa with I outer bristle at middle. Mid and hind femora with I preapical anterior bristle. Fore tibia with I ad at apex; mid tibia w ith 2 ad and 2 pd; hind tibia w ith I ad at base and 2 pd at apex. Relative lengths of segments of legs LI 2.2 : 1.0: 0.7: 0.5: 0.3: 0.3 ; LII 2 .7: 1.6: 1.2: 0.8: 0.6: 0.3; LIII 3.2: 0.7: 1.4: 0.9: 0.7: 0.3 . Wing slightly tinged with grey; ve ins dark brown, R4+5 and M sl ightly convergent apically; CuAx ratio 0.43. Squama dark brown, with black ha irs. Halter dark brown.

Abdomen meta llic green with grey po llen. Hairs and bristl es on abdomen black. Male genitalia (Fig. 48): Epandrium with long finger- like lateral lobe bearing I th ick apical bristle; cercus with short or long ha irs; ae­deagus trifurcate

Female. Unknown.

Holotype male, Sichuan : Emei , Jingshui (650 m), 1998. VIII. I 0 , Ding Y ANG . Paratype 1 male, Sichuan: Qing­cheng Mountain, 1998. VIII. 22, Ding YANG. (CAU & SKU)

Distribution. China (Sichuan) .

Etymology. The species is named after the aedeagus trifurcate apica lly .

Remarks. The new species is easily separated from other known species of the genus from China by having the R2+3 and R4+5 distinctly th ickened basally.

Subfamily Sympycninae

27. Teuclzopltorus emeiensis sp. nov. (Figs. 49-51 )

Diagnosis. Mid femur with a row of I 1-13 pv on apical half. Fore tarsomeres 2-4 distinctly shortened.

Male. Body length I .3- I .4 mm, wing length I .5- I .6 111111.

Head metallic green with grey pollen. Eyes contiguous on face . Hairs and bristles on head black; postocular bristles (including ventral hairs) black. Ocellar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna black; first flagellomere nearly triangular, as long as wide; arista black, with very short basal seg­ment. Proboscis dark brown, with black hairs; palpus black, with black hairs and I black bristle.

Thorax metallic green with grey pollen. Hairs and bristles on thorax black; 5 strong de, 6-7 uniseriate acr somewhat long; I h, I ph, I ih, I su, 2 npl, 2 sa, I pa; scutellum with two pairs of bristles (outer pair very short, hair-like) and several very short marginal hairs. Legs black. Hairs and bristles on legs black; fore coxa with 6 apical bristles; mid coxa w ithout outer bristle; hind coxa with I outer bristle near base. Fore femur with 4 pv apical Iy; mid femur with a row of I 1-13 pv on apical half; hind femur w ith I long curved av. Fore tibia with a row of short thick av; mid tibia with 2 ad and I pd; hind tibia w ith 2 ad, 4 pd (of which 3 bristles are located apically) and a row of 8 v (of which 3 apical bristles are thick) . Relative lengths of segments of legs LI 1.4: 0.6: 0.25: 0.2 : 0.2: 0.3; LII 2.2: 0.75: 0.4: 0.35 : 0.3: 0.3; LIII 2.3: 0 .5: 0.6: 0.4: 0.3 : 0.3 . Wing tinged with brown; veins dark brown, R4+5 and M slightly divergent api­cally; costal cal lus absent; CuAx ratio 0.4. Squama dark brown, with black hairs. Halter dark brown.

Abdomen metallic green with grey pollen. Hairs and by.stles on abdomen black. Male genita lia (Figs. 49-50): /~pandrium with distinct lateral lobe .bearing conical pro­cess basally, surstylus nearly conical apically; cercus rather short and obtuse apically.

Female. Unknown.

Holotype male, paratype I male, Sichuan: Emei Moun­tain, Jingshui (700-750 m), I 998. VIII . 15, Ding YANG. Paratype I male, Sichuan: Emei Mountain, Jingshui (650 m), 1998. VIII. 10, Ding Y ANG. (CAU & SKU)

Distribution. China (Sichuan).

Etyrnology. The species is named after the type local ity, Emei Mountain.

Remarks. The new species is somewhat similar to the ventralis sp. nov. in that the eyes are contiguous on the face and that the epandrium has a latera l lobe, but may be easily separated from the latter by the short first flagel lo­mere.

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240 by Ding YANG and Toyohei SAIGUSA

... . . '· ·;:.;,-:, -- I I

I /

I

/ /

49

----- ......

/,- -;> I , '

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I I I I

I I ,---- ... _ / /

.......... /.... --__ ..... ..-, ... .; I ~------I I I

I I I I

51

50

Figs. 49-51 - Teuchophorus emeiensis, sp. nov. , male. 49. surstylus and lateral lobe on epandrium, ventral view; 50. genitalia,

lateral view; 5 1. antenna (excluding scape and pedicel), lateral view.

I L-,

I /

........... - 1

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54

Figs. 52-54 - Teuchophorus ventralis. sp. nov., male. 52. hind femur and tibia, lateral view; 53. antenna (excluding scape and

pedicel), lateral view; 54. geni tali a, lateral view.

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Dolichopodidae from China (V III ): Emei moutain • 24 1

28. Teuchophorus ventralis sp. nov. (Figs. 52-54)

Diagnosis. First fl age ll omere much elongated and thin apica lly. Mid and hind femora with 1-2 v. Fore tarsomere I as long as rest of fo re tarsus.

Male. Body length 1.5-1.6 mm, wing length 1.7-1 .8 mm. Head metalli c green with grey pollen. Eyes contiguous

on face. Hairs and bristles on head black; postocular bristles (including ventra l hairs) black. Ocell ar tubercle weak, with one pair of long oc and 2 very short posterior hairs. Antenna bl ack; first flage llomere much elongated and thin apically; ari sta bl ack, bare except apex very short pubescent, with very short basa l segment. Proboscis dark brown, with black hairs; palpus bl ack, with bl ack hairs and I black bri stl e.

Thorax metallic green with grey poll en. Hairs and bri stl es on thorax bl ack; 6 strong de, 7 uniseriate acr somewhat long; I h, I ph, I ih , I su, 2 npl , 2 sa, I pa; scutellum with two pairs of bristles (outer pair very short, hair-like) and several ve ry short marginal hairs. Propleuron with I short hair on upper portion and 1-3 short hairs on lower portion. Legs bl ackish. Hairs and bristles on legs black; fo re coxa with 6 lateral and api cal bri stl es; mid coxa without outer bri stl e; hind coxa with I outer bristle near base. Mid femur with I spine-like vat basa l 1/3, .apica lly with 3 pv; hind fe mur with 2 v at middle and I long av apica ll y. Fore tibi a weakly curved, without di stinct d; mid tibi a basa lly with I ad and I pd; hind tibia sli ghtly curved and weakly thi ckened apically, with two rows of thin v (av longer basa lly, pv longer apically). Fore tarsomere I as long as fore tarsomeres 2-5 together, basa lly with thi ckened ventral surface bearing a row of 5 spine-like ventral bristles. Relati ve lengths of segments of legs LI 1.7: 1.3: 0.2: 0.3: 0.35: 0.3 5; LII 2.7: 1.2: 0.5: 0.4 : 0.3: 0.3; LIII 3. 1: 0.7: 0.8: 0.4: 0.3: 0.3. Wing hya line; ve ins dark brown, R4+5 and M parallel apically; costal callus ab­sent; CuAx ratio 0.4. Squama dark brown, with black hairs. Halter black.

Abdomen metalli c green with grey poll en. Hairs and bri stles on abdomen bl ack. Male genitalia (Fig. 54): Epandrium with lateral lobe bearing conica l process ba­sa lly, surstylus nearl y conica l api cally; cercus rather short and obtuse api ca ll y.

Female. Body length 1.5-1. 8 mm, wing length 1.7-1.8 mm. Simil ar to male, but legs somewhat pale and normal. Mid tibi a with 2 ad, I pd and I av.

f-lolotype male, paratypes 26 males 40 females, Sichuan: Emei Mountain , Jingshui (700-750 m), 1998. Vlll. 15, Ding YANG. Paratype I male, Sichuan: Emei Mountain, Wuxiangang (700 m), 1998. VIII. 14, Ding YANG; 3 males, Sichuan: Emei Moun ta in, Jingshui (65 0 m), 1998. VIII. 10, Ding YA G. (CAU, SKU & IRSN)

Distribution. China (Sichuan).

Etymology. The species is named after the mid and hind femora with spine-like ventral bristles.

Remarks . The new species is eas ily separated from other known species of the genus from China by the first fl age llomere much elongated.

29. Teuclwplwrus sinensis Y ANG et SAIGUSA

Teuchophorus sinensis YANG et SAIGUSA, 2000. In : In­sects of the Funiu Mountains region (2). Type locality: Henan.

Specimen examined. I male, Sichuan: Emei Mountain, Jingshui (650 m), 1998. Vfll. I 0, T. SAIGUSA (SKU).

Distribution. China (Sichuan, Henan).

Acknowledgments

Our sincere thanks are due to Prof. H. SHIMA and Assoc . Prof. 0. YATA (Kyushu University, Fukuoka), Mr. K. MASUNAGA (Lake Biwa Mu­seum. Shiga), and Dr.. P. GROOTAE RT (Roya l Be lgian Institute of Natural Sciences, Brussels) for their kind help in many ways. The resea rch is part ia lly supported by the Japan Society fo r the Promotion of Science under Postdoc tora l Fel lowships fo r Foreign Researchers awarded to the fi rst author.

The present paper is cont ri bution No. 60 from the Biosystematic Laboratory, Graduate school of Soc ial and cul tural studies, Kyushu Uni vers ity.

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Ding YANG* Biosystematics Laboratmy

Graduate School of Social and Cultural Studies Kyushu University

Ropponmatsu, Fukuoka, 810-8560 Japan

T. SAIGUSA 7- 1-402, Baikoen 2-Chome, Fukuoka, 810-0035 Japan

(*) Permanent address: Department of Entomology China Agricultural University Haidian, Beijing I 00094, China)

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